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Suborder Trogiomorpha Suborder Troctomorpha Suborder Psocomorpha

Infraorder Atropetae


Subfamily Thylacellinae

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Thylacella cubana (Banks) 1941

Thylacella cubana map. Thylacella cubana female.   Size:2.5 mm. Range: This is the only species of the genus in the Western Hemisphere. Known only from females. It has been colleted around Brownsville, TX and penninsular Florida. Outside the U.S. it occurs in Cuba, Mexico, Guatemala and Belize. Habitat: persistant dead leaves of various plants,espcially palms, also on trunks and branches of trees.

Subfamily Lepidopsocinae

Echmepteryx hageni (Packard) 1870

Echmepteryx hageni map echmepteryx hageni female   Size: ~ 3 mm. Range outside Texas: Eastern U.S. west to Arkansas Habitat: Bark and leaves of deciduous trees and on exposed outcrops. Males are rare except at a few sites in the U.S.

Subfamily Echinopsocinae

Neolepolepis occidentalis (Mockford) 1955

Neolepolepis occidentalis map. Neolepolepis occidentalis female   Size: ~1.6 mm. Range outside Texas: Eastern U.S.; less common in the southwest. Known from both sexes. Habitat: Leaf litter and dead hanging leaves. There is a macropterous form of this species and I would be delighted to encounter it.


Cerobasis guestfalica (Kolbe) 1880

Cerobasis guestfalica female   Size:1.5 mm. Cosmopolitan. Habitat: Deciduous and evergreen trees and stone outcrops. Identified by the anchor shaped mark on the frons. Most populations are parthenogenetic.


Lepinotus reticulatus Enderlein 1905

Lepinotus reticulatus female   Size: ~ 1.2 mm. Range: Cosmopolitan. Habitat: Leaf litter, granaries, bird and mammal nests. Winglets with a distinctive reticulate pattern. The wings are easily dislodged. This obligate parthenogen can become a pest in stored food products.

Trogium pulsatorium (Linnaeus) 1758

Trogium pulsatorium map Trogium pulsatorium   Size: ~ 1.7 mm. Range:  Not common; found primarily in the northeast; Habitat: Can occur in grain mills, bird nests, even herbaria. I have only found it on palms (Sabal minor) at Aquarena Springs. Males and females look very much alike.


Rhyopsocus bentonae Sommerman 1956

Rhyopsocus bentonea map Rhyopsocus bentonea   Size: male 1.03 mm, female 1.18 mm Range outside Texas: throughout penninsular Florida, Atlantic Coast to Brunswick GA, around Gulf Coast to Texas, southern and southeastern Mexico. Habitat: dead leaves of palm,yucca and Typha. In this photo the male is on the left. I call this one the "Hobbit psocid".

Rhyopsocus eclipticus Hagen 1876

Rhyopsocus eclipticus map Rhyopsocus eclipticus female Rhyopsocus eclipticus male   Size: 1.4 mm. Range: Atlantic Coastal Plain, Gulf Coast from Florida to south Texas, along Mississippi embankment to southern Illinois. Habitat: dead persistent leaves, conifer foilage and leaf litter. Note the large terminal segment of the maxillary palps.



Dorypteryx domestica (Smithers) 1958

Dorypteryx domestica   Size: ~ 2 mm. Range outside Texas: records from central Europe, southern England, Zimbabwe. It is only known from domestic situations such as basements. This rather odd looking psocid reminds me of Gollum from Lord of the Rings.

Image by Patrick-Emmanual Boulanger-Nadeau.

Psocathropos lachlani (Enderlein) 1903

**** ****   Size: ~ 1.5 mm. Range:Cosmopolitan. It has been recorded in Florida and Texas. Habitat: Often found in buildings, north-facing outer walls of buildings and on natural vegitation. This is a sexual species. The sex of the specimen shown is not known.

Image by Joseph Connors.

Psyllipsocus apache Mockford 2011

Psyllipsocus apache map Psyllipsocus apache female Psyllipsocus apache male   Size: 1.2 - 1.5 mm. Range outside Texas: Records from Arizona and California. Habitat: Found in leaf litter, dead yucca leaves and dead palm leaves. All are brachypterous. Named for Indian tribe from type locality.

Psyllipsocus decoratus Mockford 2011

****   Size: ~ 3 mm. Range: There are records from two sites in Arizona and two sites in Texas. Habitat: Rock outcrops near cave entrances. John Schneider recorded this species on brick walls with a patina of algae and lichens in Houston, TX. This species was described from nymphs collected near a cave entrance in Cochise Co.AZ. The sex of the specimen shown is unknown.

Image by Turner Brockman.

Psyllipsocus ramburii Selys-Longchamps 1872

macropterous female Psyllipsocus ramburii female   Size: ~2 mm Range: cosmopolitan. Females micropterous or macro- pterous. Found in caves, cellars and occasionally shaded outcrops. Records:Texas Memorial Museum.

Micropterous image by Matthew Bergeron; macropterous one by Stepen Luk.

Suborder Troctomorpha

Infraorder Amphientometae


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Lithoseopsis hellmani (Mockford & Gurney) 1956

Lithoseopsis hellmani map Lithoseopsis hellmani female   Size:3.5 mm Range outide Texas: southern Arizona south to Central America. - Habitat: Shaded limestone outcrops. This species is part of a complex of closely related species found from southern Texas. Known from female only.

Stimulopalpus japonicus Enderlein 1906

**** ****   Size: ~ 5 mm. Range outside Texas:This species is known from the eastern U.S., Hawaii, Southeast Asia, and India. Habitat: Stone outcrops as well as cement structures with a patina of algae and lichen. This is a parthenogenetic species.

Image by Ashley Bradford.

Infraorder Nanopsocetae


Subfamily Embidopsocinae

Belaphotroctes sp.

Belaphotroctes female Yes, there are liposcelidids with wings! In this genus mx4 is much wider than mx3. Females may be apteous or macropterous depending on species.

  Size:1.7 mm Range: Winged females have been found in Hays Co. This specimen was caught sweeping grass. This one is a possible ant mimic. We hope to find apterous males and females of this species.


Embidopsocus laticeps Mockford 1963

Embidopsocus laticeps   Size: ~ 1.0 mm Range: Texas- Harris Co.; Gulf Coast Florida to Texas and in Bullock Co. Ga. Habitat: under bark of dead trees. Known from males, macropterors and apterous females.

Photoghaph by Graham Montgomery.

Subfamily Liposcelidinae

Liposcelis Section I

Liposcelis brunnea Motschulsky 1852

brunnea   Size: ~ 1.1 mm Range: west Texas, Arizona, New Mexico, Colorado, northeast toOttowa, Ontario. Habitat: Conifer and oak foilage, and ground litter beneath these trees, bird nests and domestic situations.

Image by George Opit.

Liposcelis deltachi Sommerman 1957

Liposcelis deltachi   Size: 1.1 mm Range: Hays Co.; Southern and western Texas, southern New Mexico, southern Oklahoma; San Luis Potosi, Mexico. Habitat: Foliage of junipers, pine, yuccas, mesquite and leaf litter. Known from females only.

Liposcelis entomophila (Enderlein) 1907

Liposcelis entomophila   Size: ~ 1.1 mm. Range: Texas-Hays Co.;  Midwest and southeastern U.S. to central Texas; fairly cosmopolitan. Habitat: Primarily domestic but also taken in leaf litter. Field collected specimens (as in leaf litter) mostly tropical and subtropical. Known from male and female.

Liposcelis fusciceps Badonnel 1968

**** ****   Size: ~ 1.2 mm Range: Texas Hays Co, and in Arizona Habitat: grasses and dead vines. Known from female only. Collected in Brazil in the nest of an ant, Camponotus rufipes. Dorsal view on the left and ventral on the right. Note "Head medium brown with a slight reddish tinge: palpi colorless".

Liposcelis hirsutoides Mockford 1978

Liposcelis hirsutoide   Size: 1.1 mm. Range: central and south Texas and in central-penninsular Florida.  Habitat: woody vegetation and under bark. I find it especially on lichen covered banches of Mesquite.  Known from both sexes but I have yet to photograph a male.


Liposcelis ornata Mockford 1978

Liposcelis ornata female   Size: 1.2 mm. Range: Gulf Coast from Florida to Texas and in the Mexican states of San Luis Potosi and Tabasco. Known from female only. Habitat: Great variety of trees and shrubs. I have found in in leaf litter and dead palm fronds.

Liposcelis pearmani Lienhard 1990

Liposcelis pearmani   Size: ~ 1.1 mm. Range: Texas- Hays Co.  Relatively cosmopolitan; recorded from several U. S. states including Texas. Habitat: Domestic; also found in bird nests, under bark and sweeping grass. This species is known from both sexes.

Liposcelis pallida Mockford 1978

Liposcelis pallida   Size: ~ 1 mm. Range: Texas- Blanco Co.; Known from the Davis Mts. of Texas and Catalina Mts. of Arizona. The specimen pictured was collected at Flat Creek (Blanco Co.). Habitat: Dead persistent leaves of yucca; leaf litter. Known from both sexes.

Liposcelis Section II

Liposcelis bostrychophila Badonnel 1931

Liposcelis bostrychophila female   Size: ~ 1.0 mm Range: Texas- Hays Co.;  Cosmopolitan Habitat: under bark, bird nests, stored grain, old books, chicken litter. Known only from females. If there is a classic booklouse, this is it.



Nanopsocus oceanicus Pearman 1928

Nanopsocus oceanicus map Nanopsocus oceanicus female Nanopsocus oceanicus male   Size: ~ macropterous female, 1.5 mm., apteous female 1.0 mm. Range: southeastern U. S. north to Illinois; also Mexico, Central America, West Indies, West and Central Africa, Japan and South Pacific Islands . Habitat: Occurs on dead leaves of palms, bamboos and grasses.

See frame below for macropterous female.

Nanopsocus oceanicus mac female

Nanopsocus oceanicus mac female Nanopsocus oceanicus mac female   Macropterous females brown; apterous females creamy white; males (apterous) creamy white but with a reddish brown stripe along anterior margin of abdominal terga 2-7. This genus is characterized by a foliate shaped hind tarsal claw on each foot.

Peritroctes bengalensis Thornton 1966

Peritroctes bengalensis map Peritroctes bengalensis female   Size: 1.1 mm. Range: First recorded from the Botanical Gardens of Calcutta, India by Ian W. B. Thornton and S. K. Wong, "Some Psocoptera from West Bengal, India", Transactions of the Royal Entomological Society of London, 118(1):1-21, 1966. Habitat: leaf litter, dead palm frinds, dead persistent leaves.

Tapinella maculata Mockford and Gurney 1956

Tapinella maculata map Tapinella maculata male Tapinella maculata female   Size: male ~ 1 mm, female ~ 1.5 mm. Range: Central Texas to lower Rio Grande Valley, Mexico and South America. Habitat: dead palm and grass leaves Known from males, macropterous and apterous females.

See frame below for macropterous female.

Tapinella maculata... continued

Tapinella maculata female Tapinella maculata male   

The macropterous female (length 1.7 mm) is, in my experience, rarer than the apterous one. In this family, the wings are held flat over the abdomen, not roof-like as in the Psocomorpha. Dr. Mockford says that this individual from Gonzales Co. shows much more patterning on the head than specimens in far south Texas.

Tapinella sp. "Type 2"

Tapinella sp. map Tapinella sp. female Tapinella sp. male   Size: male ~ 1.1 mm, female ~ 1.3 mm Range: unknown Habitat: Leaf litter This un-described species lacks the backwards "U" shaped marks along the lateral edge of the abdomen. This species often occurs with Tapinella maculata.

See frame below for macropterous female.
Tapinella sp. Type 2 Tapinella sp. Type 2   

The images on this row depict the macropterous form of Tapinella sp. Type 2. These individuals possess ocelli while the apterous morphs do not.


Sphaeropsocus bicolor Mockford 2013

Sphaeropsocus bicolor map Sphaeropsocus bicolor   Size:0.7 mm Range: single site Habitat: leaf litter This species has not been found in Texas but it is so remarkable that I had to include it. This remarkable species was discovered by Tim King. "It is the first living species of its genus, known otherwise only from Sphaeropsocus kuenowii Hagen, a fossil species from the Baltic amber (Eocene)." Mockford, E.L., A New species of Sphaeropsocus Hagen from south eastern U. S.: the first living species of its genus, Life: the Excitement of Biology 1(2): 100-111.

Suborder Psocomorpha

Infraorder Archipsocetae


Archipsocus sp.

Archipsocus sp.   Archipsocus specimens are found along the Gulf coast from Florida to Texas and in Mexico. They can occur in Texas as far inland as Travis Co. I have not yet identified this species. These specimens were found at Palmetto State Park. There are three described species which might occur on the Texas coast: Archipsocus floridanus, A. gurneyi A. nomas. The webbing from these psocids often covers entire tree trunks and causes a media sensation.

Epipsocetae   Caeciliusetae   Homilopsocidea 1   Homilopsocidea 2   Psocetae

Infraorder Caeciliusetae

Caeciliusidae   Stenopsocidae   Amphipsocidae   Dasydemellidae



Polypsocus corruptus (Hagen) 1861

Polypsocus corruptus map Polypsocus corruptus female Polypsocus corruptus male   Size: ~ 3.5 mm  Range outside Texas: eastern U. S. west to Missouri; eastern Canada and Pacific coast from British Columbia south to northern Califorina. There are closely related species in Mexico. Habitat: Variety of broad-leaf  trees including Magnolia.


Stenocaecilius antillanus (Banks) 1938

****   Size: 2.8 mm Range: From south Florida, south Texas, Mexico, Surinam and Cuba Habitat: Found on dead foliage of palms, grasses and cycads This species is parthenogenetic. This specimen was collected in Kingsville, TX.

Stenocaecilius casarum (Badonnel) 1931

Stenocaecilius casarum map Stenocaecilius casarum female   Size: ~ 3.5 mm Range: outside Texas- Gulf Coast from south Florida to the Rio Grande Valley. Wide range in the tropics including Mexico, Guatemala & Venezuela. Habitat: Living or dead palm foilage. With the exception of a single male taken in Guyana, this species consists only of females.

Valenzuela croesus (Chapman) 1930

Valenzuela croesus map Valenzuela croesus female Valenzuela croesus male   Size:~ 3 Range: Texas, Arkansas east to Mississippi, North Carolina and north the New York; ranges from Mexico to Guatemala. Habitat: Prefers foilage of Juniperus and Cupressus.


Valenzuela flavidus (Stephens) 1836

Valenzuela flavidus Valenzuela flavidus female Valenzuela flavidus male   Size: 3.6 mm, Range outside Texas: eastern U. S., northern Pacific Coast, also Europe, Canary Islands, Bermuda, and Mexico. Habitat: Foilage of broadleaf trees. Males are scarce throughout most of the range. Populations which have males may represent a sibling species (Mockford, 1993).

Valenzuela indicator Mockford 1969

Valenzuela indicator map Valenzuela indicator female Valenzuela indicator male   Size: ~ 4 mm, Range outside Texas: Florida, Georgia, Cuba, Habitat: found on various trees including oaks, palms and on Sapnish moss.

Images by Robert Zimlich.

Valenzuela lochloosae (Mockford) 1965

Valenzuela lochloosae map Valenzuela lochloosae female Valenzuela lochloosae male   Size: ~ 3 mm Range: Throughout Florida and southeast Alabama. Range probably along entire Gulf Coast. Habitat: persistent dead leaves of native grasses.

Valenzuela manteri (Sommerman) 1943

Valenzuela manteri map Valenzuela manteri female Valenzuela manteri female   Size: ~ 3 mm Range outside Texas: eastern U. S. records in Minnesota, Missouri and Nebraska. Habitat: dead leaves of cattail, corn and palmetto. This species was described by Kathryn M. Sommerman in "Description and bionomics of Caecilius manteri", Proc. Ent. Soc. of Washington, 45(2): 9-39, 1943 with marvelous illustrations. Males are rare in this species.

Valenzuela micanopi (Mockford) 1965

Valenzuela micanopi map Valenzuela micanopi female Valenzuela micanopi male   Size: male 3.0 mm, female 3.2 mm Range outside Texas: throughout Florida, southeast Alabama and on Bimini Island and the Bahamas. Habitat: dead leaves of Sabal and Coccothrinax.


Valenzuela nadleri Mockford 1966

Valenzuela nadleri map Valenzuela nadleri female Valenzuela nadleri male   Size: male 2.6 mm, female 2.7 mm. Range: eastern US, west to Wisconsin and Missouri, also SE Canada. Recent records extend range westward. Habitat: leaf litter. Females dimorphic- macropterous and micropterous.

See frame below for micropterous female.
Valenzuela nadleri micro female   This is an image of the micropterous female by Jean Brodeur of Canada. This female morph looks very similar to the micropterous female of Valenzuela posticus. Micropterous females of V. nadleri, however, have dark antennae all the way to the tip. Both  occur in leaf litte

Valenzuela posticus Banks 1914

Valenzuela posticus map v. posticus female v. posticus male   Size:male 3-4 mm, micropterous female 2 mm. Range outside Texas- records from Florida, Illinois, Michigan and New York. Females either fully winged or microptreous. The micropterous V. nadleri which looks very similar has uniformly dark antennae while those of V. posticus are pale terminally.

Male image by Ken Allen, female image by John F. Carr.

Valenzuela subflavus (Aaron) 1886

subflavus map Valenzuela subflavus female Valenzuela subflavus male   Size: ~ 2.5 mm  Range: Outside Texas - Atlantic and Gulf Coast  Habitat:  foliage of broadleaf evergreen trees like American holly and live oak.


Teliapsocus conterminus (Walsh) 1863

Teliapsocus conterminus Teliapsocus conterminus female Teliapsocus conterminus male   Size: 5.6 mm Range: Throughout most of U.S. and Canada, but absent from northern midwestern states. Habitat: Great variety of broad-leaf and coniferous trees and occasionally ground litter. Not known outside North America.

Male image by Patrick Coin.


Graphopsocus cruciatus (Linnaeus) 1768

Graphopsocus cruciatus map Graphopsocus cruciatus female Graphopsocus cruciatus male   Size:  ~3.8 mm   Range:  Texas Gulf Coast. This non-native species was apparently introduced along both coasts sometime in the early 19th century. Habitat: Foilage of braodleaf deciduous and evergreen trees.

Infraorder Epipsocetae


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Bertkauia crosbyana Chapman 1930

Bertkauia crosbyana map Bertkauia crosbyana female   Size: ~ 1.7 mm  Range: throughout eastern U.S., southern Rocky Mts., northwest Washington state. Mexico: San Cristobal and Chiapas. Habitat: woodland ground litter Males (very rare) macropterous, femaes micropterous.

A Disturbance in the Force …

Within the suborder Psocomorpha, the infraorder Homilopsocidea has been a problem. Molecular generic studies have cast doubt on the monophyly of this taxon. A recent elegant study by Oscar Fernando Saenz Manchola, Alfonso Gracía Aldrete, Kevin P. Johnson, and three other authors splits this infraorder into two monophyletic groups: Homilopsocidea 1 (Ectopsocidae and Peripsocidae) and Homilopsocidea 2 (Elipsocidae, Lachesillidae, and Mesopsocidae). I will follow this taxonomic change here.

Homilopsocidea 1

Ectopsocidae   Peripsocidae  


Ectopsocopsis cryptomeriae (Enderlein) 1907

Ectopsocopsis cryptomeriae map Ectopsocopsis cryptomeriae female Ectopsocopsis cryptomeriae male   Size: ~ 1.7 mm Range: Outside Texas- Gulf States from Texas to Florida and north to New York, west to Illinois Habitat: leaf litter; hanging dead leaves.


Ectopsocus californicus (Banks) 1903

Ectopsocus californicus map Ectopsocus californicus female Ectopsocus californicus male   Size: ~ 2.4 Range: Outside Texas: U. S. Atlantic and Pacific Coast, Mexico and the highlands of Guatemala Habitat: living and dead foliage of broadleaf and coniferous trees.

Ectopsocus meridionalis Ribaga 1904

Ectopsocus meridionalis map Ectopsocus meridionalis   Size:2.4 mm Range outsie Texas: throughout eastern U.S. south to Florida; central Mexico to Chile, southern Europe, Africa and Japan. Habitat: persistent dead leaves. Known only from female.

Ectopsocus richardsi Pearman 1929

**** Erichardsi   Size: 1.2-1.5 mm Range: Cosmopolitan with records in the U.S. from south Florida and south Texas. Habitat: Generally associated with stored food products, but also found in natural areas.. This species exhibits some interesting behavior documented on my YouTube channel "Diane Young Psocoptera"

Ectopsocus vachoni Badonnel 1945

Ectopsocus vachoni map Ectopsocus vachoni female Ectopsocus vachoni female   Size: males ~ 1.0 mm, females ~ 1.7 mm Range: Outside Texas-US Gulf States, California; Mexico, South America, and Europe Habitat: leaf litter, dried grass and lower foliage of trees.

See frame below for micropterous female'.
Ectopsocus vachoni female   In this species, the males are micropterous and the females are either macropterous or micropterous. The tiny male has a dark clunial comb (visible in a dissecting microscope) which identifies it as mature. I once observed a male approaching a female and vibrating his body rhythmically as a prelude to courtship. This species is likely very common throughout Texas.


Peripsocus madidus (Hagen) 1861

Peripsocus madidus map Peripsocus madidus female Peripsocus madidus male Peripsocus madidus brachypeterous   Size: ~ 3.2 mm Range: Outside Texas, Nova Scotia south to Florida and west to central Texas Habitat: bark and branches of broad-leaf and coniferous trees.

The barchypterous form of this species is shown below. I do not find this form in central Texas but did record them from Austin Co.


Peripsocus stagnivagus Chapman 1930

**** ****   Size: 1.7-2.1 mm. Range: Eastern North America and Europe. Habitat: Trunks and branches of trees, Spanish moss and dead palm fronds. Most populations are parthenogenstic except for areas in Illimois and Texas.

Peripsocus subfasciatus (Rambur) 1842

Peripsocus subfasciatus Peripsocus subfasciatus   Size: ~ 2.7 Range: Outside Texas - eastern U.S., west to Minnesota and Arkansas, Pacific Coast from San Francisco Bay to British Columbia. Habitat: branches of broadleaf and coniferous trees and shaded stone outcrops. Male virtually absent except in Washington state and Vancouver, British Columbia.

Peripsocus minimus Mockford 1971

Peripsocus minimus map Peripsocus minimusfemale Peripsocus minimus male   Size: ~ 2.0 mm  Range:  Outside Texas- southern Illinois southwestward to Missoiri, Arkansas and central Texas  Habitat: foliage of Juniperus virginiana and Juniperus ashei.

Peripsocus sp.

Peripsocus sp. mystery map Peripsocus sp female Peripsocus close to stagnivagus male   Size: male: ~ 2.2 mm, female ~ 1.5 mm Range: Habitat: Lichen covered branches of Quercus virginiana and Texas persimmon (Diospyros texana). This is an un-described species in which the female is micropterous.

Infraorder Homilopsocidea 2



Andra Group

Lachesilla kola Sommerman 1946

kola map Lachesilla ola female Lachesilla kola male   Size: ~ 2.2 mm  Range: Outside Texas- Records from southern California, Arizona and New Mexico  Habitat: specimens shown found on dead Chinaberry branches near the Concho River.

Lachesilla nubilis (Aaron) 1886

Lachesilla nubilis map Lachesilla nubilis female Lachesilla nubilis male   Size: ~ 2.6 Range: Outside Texas- throughout eastern U.S. west to New Mexico, Colorado and Utah.  Habitat: dead grasses and dead limbs with persistent leaves.

Lachesilla texana Mockford & Garcia Aldrete 2010

Lachesilla texana map Lachesilla texana female Lachesilla texana male   Size: ~ 2.2 Range: Outside Texas: Mexico: Nuevo Leon, Coahuila, and Cerrode la Silla. - a second population in the Smokey Mts. Habitat: dead leaves on clumps of native grasses.

Corona Group


Lachesilla michiliensis Garcia Aldrete 1991

Lachesilla michiliensis map Lachesilla michiliensis female   Size: ~ 1.5 mm Range: Outside Texas  Mexico: Durango State Habitat: under loose bark Known only from female. "This species is neotenic as indicated by the brachyptery, small oceli and almost complete absence of sensory fields on the paraprocts" Garcia Aldrete, A., "Lachesillidae from the Biosphere Reserve 'La Michilia", Durango and Surrounding Areas, Folia Entomologica Mexicana 81: 165-183, 1991.

photograph by Graham Montgomery

Forcepeta Group

Lachesilla anna Sommerman 1946

Lachesilla anna map Lachesilla anna female Lachesilla anna male   Size:3.2 mm Range: Outside Texas occurs throughout eastern U.S. and southeastern Canada, south to Marian Co. Florida and west to Texas Co. Missouri. Habitat: persistent dead leaves trees and shrubs.

Lachesilla bottimeri Mockford & Gurney 1956

Lachesilla bottimeri map Lachesilla bottimeri waiting for image of male   Size: 2.2 mm Range: In Texas, restricted to lower Rio Grande valley; also occurs in a long north south distribution in eastern Mexico. Habitat: Found mostly on dead hanging leaves of palms. This is the only species in the Forcepeta Group in which the claspers in the male point inward.

Lachesilla contraforcepeta Chapman 1930

Lachesilla contraforcepeta map Lachesilla contraforcepeta female Lachesilla contraforcepeta male   Size: ~ 2.0 Range: Outside Texas- throughout eastern U.S., and Rocky Mountain region, New Mexico, Colorado, Idaho. Habitat: conifers such as junipers and pine. Male claspers curve outward.

Lachesilla floridana Garcia Aldrete 1999

Lachesilla floridana map Lachesilla floridana female Lachesilla floridana male   Size: 2.0 mm  Range: Outside Texas coastal distribution in Florida and Alabama;Habitat: Found on dead palm and palmetto fronds. Note the crossvein between the pterostigma and vein Rs.

Lachesilla forcepeta Chapman 1930

Lachesilla forcepeta map Lachesilla forcepeta female Lachesilla forcepeta male   Size: ~ 2.0 mm Range: Outside of Texas- throughout eastern U.S. west to Oklahoma and Kansas and the San Antonio area of Texas. Habitat: conifers including Juniperus ashei.


Lachesilla kathrynae Mockford & Gurney 1956

Lachesilla kathrynae map Lachesilla kathrynae female Lachesilla kathrynae male   Size: ~ 2.5 mm Range: Lower Rio Grande Valley; Mexico: Tabasco State Habitat: dead foliage of palms and carrizo grass.

Lachesilla penta Sommerman 1946

Lachesilla penta map Lachesilla penta female Lachesilla penta male   Size: ~ 2.5 Range: Outside Texas- Gulf states from Texas to Florida; eastern Mexico south to Chiapas State. Habitat: prsistant dead leaves of deciduous trees and palms.

Pedicularia Group

Lachesilla otomi Mockford 2002

L otomi   Size: 2 mm Range: From coastal Florida and Texas to Mexico and Central America.Habitat: Dead hanging leaves of native grasses, Palmetto, and various shrubs.

Lachesilla pedicularia (Linnaeus) 1758

Lachesilla pedicularia map Lachesilla pedicularia female Awaiting image of male   Size: 2.5 mm. Range: This species has been recorded from most of the U.S., nearly cosmopolitan elsewhere. Habitat: Found on grasses, dead persistent leaves, dried grain and occcasionally conifers. It is easily transported in human commerce and can become a pest of stored grain.

Lachesilla rena Sommerman 1946

Lachesilla rena map Lachesilla rena female Awaiting image of male   Size:1.8; Range: Outside Texas - southern California and Maricopa Co. Arizona and throughout much of Mexico. Habitat: dead grasses and low vegetation.

Lachesilla tectorum Badonnel 1931

Lachesilla tectorum map Lachesilla tectorum female   Size: 2.5 mm Range: outside Texas- Gulf coast  Florida to Texas Mexico: Tamaulipas and Nuevo Leon Habitat: dead leaves of palm and tall grasses Known from female only.


Lachesilla yucateca Mockford 2002

Lachesilla yucateca map Lachesilla yucateca female Lachesilla yucateca male   Size:~ 1.7 mm Range: Outside Texas- Mexico: Oaxaca, Tamaulipas, Yucatan; Habitat: persistent dried leaves of native grasses. This species belongs to the Rena complex in which the male epiproct is extremely long and narrow.

Riegeli Group

Lachesilla tropica Garcia Aldrete 1982

Lachesilla tropica map Lachesilla tropica female Lachesilla tropica male   Size: 2.2 mm Range - Outside Texas: Florida, throughout most of Mexico, Guatemala, Honduras, Jamaica. Habitat: hanging dead leaves of trees and shrubs.

Rufa Group

Lachesilla nita Sommerman 1946

Lachesilla nita map Lachesilla nita female Lachesilla nita male   Size: 2.5 mm. Range: Gulf Coast Georgia to Texas, also in Mexico, Beliize, Guatemala, and Panama. Habitat: Found on cabbage palm, various oaks, Eleagnus and other shrubs. Note the characteristic diffuse spot in each cell from R3, R5, M1, M2, M3.

Sclera Group

Lachesilla sulcata Garcia Aldrete 1986

Lachesilla sulcata map Lachesilla sulcata female   Size: ~ 2.0 mm Range: Outside Texas- Gulf states from Texas to Florida. Mexico: Nuevo Leon, Tamaulipas, Chiapas. Habitat: native grass.The male of this species was only recently been described (Garcia Aldrete & Mockford "Reappraisal of Species Group Patzunensis" Proc. Ent. Soc. Washington 113(4): 417-425, 2011.

Infraorder Philotarsetae

Pseudocaeciliidae    Trichopsocidae   


Aaroniella badonneli (Danks) 1950

Aaroniella badonneli map Aaroniella badonneli   Size: ~ 3 mm Range: east Texas, occurs throughout much of the eastern U. S.and north to Ontario, the soviet state of Georgia and the Azores. Habitat: twigs and lichen covered branches of a variety of trees and shrubs. Known only from female.

Aaroniella sp.

Aaroniella sp. map Aaroniella sp. female Aaroniella sp. male   Size: 2.2 mm This un-described species is found at the Aransas National Wildlife Refuge. Habitat: small dead branches of live oak in a heavily wooded area.


Pseudocaecilius citricola (Ashmead) 1879

Pseudocaecilius citricola map Pseudocaecilius citricola female   Size: 2.8 mm Range: Outside Texas, occurs throughout Florida, on the Alabama coast, pan-tropical; is even recorded on the Galapagos Islands; Habitat: living leaves of citrus, evergreen oaks, palms and other trees. Known from female only.


Trichopsocus dalii (McLachlan) 1867

Trichopsocus dalii map Trichopsocus dalii female Trichopsocus dalii male   Size: ~ 2.8 mm Range: outside Texas- this introduced species is found in states on the Gulf and Atlantic coast of the U.S. and in the Atlantic and Mediteranean coast of Europe and North Africa Habitat: branches of braod leaf trees like citrus, bay, Ligustrum and Ilex.

Infraorder Psocetae


Hemipsocus chloroticus (Hagen) 1858

Hemipsocus chloroticus map Hemipsocus chloroticus female Hemipsocus chloroticusHemipsocus chloroticus male   Size: ~ 3 mm. Range outside Texas: Non native. Found South Carolina to Florida along coastal plain to Texas; Oregon as an introduction from Japan. Native to Southeast Asia and many Pacific Islands. Habitat: Forest leaf litter.


Lichenomima sparsa (Hagen) 1861

Lichenomima sparsa map Lichenomima sparsa female Lichenomima sparsa male   Size: male 4.7 mm female 5.9 mm Range: Outside Texas - Washington D.C. south to Florida, west to Indiana,Arkansas. Inhabits shaded stone outcrops and tree trunks. Dr. Mockford has expressed the need for this genus to be revised.


Lichenomima sp. "W"

Lichenomima sp. 'W' map Lichenomima sp. 'W' female Awaiting image of male   Size: female 4.8 mm Range: Outside Texas- unknown. Found on Quercus virginiana at Welder Wildlife Refuge.


Amphigerontiinae      Psocinae

Subfamily Amphigerontiinae

Amphigerontia montivaga (Chapman) 1930

Amphigerontia montivaga map Amphigerontia montivaga female Amphigerontia montivaga male   Size: ~ 4.7 mm Range: outside Texas- New York south to Florida and west to Wisconsin and south Texas Habitat: branches of broad-leaf and evergreen trees.


Blaste garciorum Mockford 1984

Blaste garciorum map Blaste garciorum female Blaste garciorum male   Size: ~ 3.5 mm Range outside Texas- south to Mexico and Honduras Habitat: branches of small trees and shrubs in arid woodlands.

Blaste longipennis (Banks) 1918

Blaste longipennis map Awaiting image of female Blaste longipennis male   Size: ~ 5 mm. Range outside Texas: Yellowstone Natonal Park south to Chiricahua Mountainsof Arizona. Habitat: Collected on dwarf mistletoe, Arceuthobuim vaginatum and conifers.

Blaste persimilis (Banks) 1908

Blaste persimilis map Blaste persimilis female Blaste persimilis male   Size: 3.5 mm. Range: Gulf Coast from Alabama to Brownsville, Texas then north to Refurio. Habitat: branches of oaks and dead vines. Known only from the male. I found females at Palm Harbor, TX. No published description yet of female.

Blaste posticata (Banks) 1905

Blaste posticata map Blaste posticata female Blaste posticata male   Size: ~ 3.8 mm Range outside Texas- south to Mexico, Honduras and Guatemala Habitat: broad-leaf trees, shrubs and pine.

Blaste quieta (Hagen) 1861

Blaste quieta map Blaste quieta female Blaste quieta male   Size: ~ 4.5 mm Range outside Texas- Montreal to Florida and west to South Dakota and Texas; outlier populations in Idaho and British Columbia Habitat: branches of broad-leaf and coniferous trees.


Blastopsocus lithinus (Chapman) 1930

Blastopsocus lithinus map Blastopsocus lithinus female Awaiting image of male   Size: ~ 3.7 Range outside Texas- Montreal south to Florida - west to Minnesota to Texas  Habitat: branches of broad-leaf and coniferus trees.

Blastopsocus walshi Mockford 2002

Blastopsocus walshi map Blastopsocus walshi female Blastopsocus walshi male   Size: ~ 3.3 mm Range outside Texas-Illinois south to Florida Habitat: branches of broad-leaf trees.

Subfamily Psocinae

Tribe Cerastipsocini

Cerastipsocus venosus (Burmeister) 1839

Cerastipsocus venosus map Cerastipsocus venosus female Cerastipsocus venosus male   Size: male ~ 6 mm female 8mm Range outside Texas - Maine south to southern Florida; south to Mexico. Habitat: Trunks and brances of broad leaf trees and conifers. Pterostigma of males is dark while those of females are white.

Images by Robert Zimlich.

Cerastipsocus trifasciatus (Provancher) 1876

Cerastipsocus trifasciatus map Cerastipsocus trifasciatus female Awaiting image of male   Size: ~ 6 mm. Range outside Texas: Southestern U.S. continuing through Arkansas to Arizona. Also in Mexico, Guatemala, Costa Rica and Columbia. Habitat: Tree trunks and branches and rock outcrops.

Image by William Geoghegan.

Tribe Metylophorini

Metylophorus novaescotiae (Walker) 1853

Metylophorus novaescotiae map Metylophorus novaescotiae female Metylophorus novaescotiae male   Size: 6-7 mm Range outside Texas: Nova Scotia and coastal Maine to Florida, west to Minnesota. Habitat: Branches of broad-leaf trees and shrubs. Also occurs in Mexico.

Images by Tom Murray.

Tribe Psocini


Hyalopsocus sp. close to floridanus

Hyalopsocus #1 map Hyalopsocus</span>
sp. female Hyalopsocus</span>
sp. female   Size: male  ~5.4 mm, female ~5.6 mm Range:   Habitat: Wide variety of braodleaf and coniferous trees; usually on dead branches encrusted with lichen.

Hyalopsocus striatus (Walker) 1853

Hyalopsocus striatus #2 map Hyalopsocus striatus 2 female Hyalopsocus striatus 2 female   Size: 5.3 Range outside Texas: Throughout northeastern U.S. and southeast Canada. Occurs spottily in western states. Habitat: Juniperous ashei  This species is characterized by the extensive pigment in cells Cu2 and IA in the forewing.

Psocus leidyi Aaron 1886

Psocus leidyi map Psocus leidyi female Psocus leidyi male   Size: ~ 5.4 mm Range: Maine to Minnesota, south to Florida and southwest to Arkansas with an outlier population in northern California. Habitat: Broadleaf trees, Mesquite, Hackberry, live oak.

Tribe Ptyctini

Indiopsocus bisignatus (Banks) 1904

Indiopsocus bisignatus map Indiopsocus bisignatus female Indiopsocus bisignatus female   Size: ~ 3.9 mm Range Outside Texas -eastern U.S. from northern Minnesota south to west Florida, west of the Mississippi it occurs spotily to Wyoming, Colorado and Texas; also occurs in Mexico Habitat: trunks and branches of braod-leaf and conifer trees.

Indiopsocus campestris (Aaron) 1886

Indiopsocus campestris map Indiopsocus campestris female Indiopsocus campestris male   Size: male 3.1 mm, female 3.2 mm Range: Atlantic Coast, throughout Florida continuing along Gulf Coast to Texas. Habitat: broad leaf trees and shrubs including live oak, turkey oak and pine.


Indiopsocus infumatus (Banks) 1907

Indiopsocus infumatus map Indiopsocus infumatus female Indiopsocus infumatus male   Size: ~ 3.6 Range Outside Texas - Oklahoma, New York, south to Kentucky; central Texas is probably at the western end of the range for this species   Habitat: dead branches of broad-leaf trees; my specimens on live oak and ashe juniper.

Indiopsocus lanceolatus Mockford & Young 2015

Indiopsocus lanceolatus map Indiopsocus lanceolatus female Indiopsocus lanceolatus male   Size: ~ 3.6 mm Range: Outside Texas: Gulf Coast, AL, GA, FL, VA. Habitat: broad leaf trees like Quercus virginiana. Recently described in Trans. Am. Entomol. Soc. 141:233-251.

Indiopsocus lacteus Mockford & Young 2015

Indiopsocus lacteus map Indiopsocus lacteus female Indiopsocus lacteus male   Size: ~ 3.9 Range: endemic to Texas Hill Country. Habitat: lower bare limbs of living Juniperus ashei. Recently described in Trans. Am. Entomol. Soc. 141:233-251.

Indiopsocus palmatus Mockford & Young 2015

Indiopsocus palmatus map Indiopsocus palmatus female Indiopsocus palmatus male   Size: ~ 3.8 Range : Apparently a Texas endemic. Habitat: braodleaf trees like Quercus virginiana, Celtis reticulata Ulmus crassifolia and Acacia farnesiana. Recently described in Trans. Am. Entomol. Soc. 141:233-251.

Indiopsocus texanus (Aaron) 1886

Indiopsocus texanus map Indiopsocus texanus female Indiopsocus texanus male   Size: ~ 3.9 mm Range outside Texas: Throughout Florida up the Atlantic coast to Virginia; Gulf Coast. Habitat: broadleaf trees including live oak.


Loensia moesta (Hagen) 1861

Loensia moesta map Loensia moesta female Loensia moesta male   Size: ~ 4.2 mm Range Outside Texas- Minnesota south to Florida west to Texas Habitat: wide variety of broad-leaf and coniferous trees.

Loensia sp. "Beauty Queen"

Loensia beauty queen map Loensia beauty queen female Loensia beauty queen male   Size: ~ 4.3 mm Range outside Texas- unknown Habitat: lichen covered branches of Mesquite. In my humble opinion, this un-described species is a beautiful psocid! Don't you agree? This species was placed (by me) in the "holding" genus Psocidus, but that genus now houses African and Asian species in which themtrichobothrial field is divided.

Ptycta polluta (Walsh) 1862

Ptycta polluta map Ptycta polluta female Ptycta polluta male   Size: ~ 3.4 mm Range: outside Texas- Nova Scotia south to Florida, west to Minnesota Habitat: branches of braod-leaf and coniferous trees.

Trichadenotecnum slossonae (Banks) 1903

Trichadenotecnum slossonae map Trichadenotecnum slossonae female Trichadenotecnum slossonae male   Size: ~ 3.2 mm Range: outside Texas- eastern U.S. west to Minnesta and central Texas Habitat: branches of broad-leaf and coniferous trees.